Abstract

Numerous vertebrates footprints have been found in the Miocene Upper Red Formation sandstone and siltstone layers in the 3 km west of Mushampa village, west Zanjan province. Most of the footprints were preserved as hypichnia, include bird’s webbed footprints Culcitapeda tridens and Culcitapeda isp.; tetradactyle footprint Antarctichnus isp. and digitigrad footprints Avipeda isp. A and Avipeda isp.B. Carnivora footprints are Bestiopeda isp. and Credontipes isp. ichnogenuses. It seems that sedimentary environment was a shallow-water pan of a marginal marine environment, and footprints formed on the exposed sediments.